When three angels are sent from heaven to protect the town of Venus Cove against the gathering forces of darkness, their mission is threatened as the youngest angel, Bethany, enrolls in high school and falls in love with another student.

In New York City, three fourteen-year-old best friends who are all daughters of celebrities watch out for each other as they try to strike a balance between ordinary high school events, such as finding a date for the homecoming dance, and family functions like walking the red carpet with their famous parents.

Told in their separate voices, sixteen-year-old Prince Oliver, who wants to break free of his fairy tale existence, and fifteen-year-old Delilah, a loner obsessed with Prince Oliver and the book in which he exists, work together to seek his freedom.

In rural Ohio, friendships and a beautiful girl prove distracting to a fifteen-year-old who has hidden on Earth for ten years waiting to develop the Legacies, or powers, he will need to rejoin the other six surviving Garde members and fight the Mogadorians who destroyed their planet, Lorien.